**Acknowledgements**

This work was supported by the Program for New Century Excellent Talents in University (grant number NCET-11-1071) of China, a NSFC (National Science Foundation of China) Grant 31260026, and a Fund for PhD in Xinjiang Production and Construction Corps (grant number 2009JC07) to W. Chen, a NSFC-Xinjiang joint Grant U1703236 to L. L. Zhang, and a Microbial Resources Utilization Innovation Team in Key Field of Xinjiang Production and Construction Crops (grant number 2017CB014) to C. X. Wan.
