**Acknowledgements**

This work was supported by Chulalongkorn University's Ratchadaphiseksomphot Endowment Fund (to Natural Biotechnology Research Unit) and Thailand National Center for Genetic Engineering and Biotechnology, National Science and Technology Development Agency (NASDA).

### **Conflict of interest statement**

The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.
