**Acknowledgements**

This work was supported by grants from the Canadian Institutes for Health Research (CIHR FDN-143345). VDM, APS, and EAM are supported by scholarships from the University of British Columbia. APS is further supported by the Frederick Banting and Charles Best Scholarship from CIHR. EAM is also supported by the Vanier Canada Graduate Scholarship from CIHR. AAG is the Canada Research Chair for Radiation Exposure Disease and this work was undertaken, in part, thanks to funding from the Canada Research Chairs program. The AAG laboratory is supported by the Canadian Institutes of Health Research.
