**Acknowledgements**

We are very thankful to Dr. Christian Capo for his help and councils regarding the redaction of the manuscript. Soraya Mezouar was supported by a "Fondation pour la Recherche Médicale" postdoctoral fellowship (reference: SPF20151234951). This work was supported by the French Government under the "Investissements d'avenir" (investments for the future) program managed by the "Agence Nationale de la Recherche" (reference: Méditerranée Infection 10-IAHU-03).
