**Acknowledgements**

The authors acknowledge the support of Fundação para a Ciência e a Tecnologia (FCT), Portugal, through grant SFRH/BD/113952/2015 and research units UID/ AGR/04129/2013 (LEAF) and UID/GEO/04035/2013 (GeoBioTec), and the Dutch Organization for Internationalization in Education (NUFFIC), through the project NICHE-MOZ-151.
