**Acknowledgements**

This work was supported by the following grants: Spanish Ministry of Economy and Competitiveness/European Regional Development Fund (MINECO/FEDER grant SAF2015-64163-R). OP is a Ramon y Cajal Fellow (RYC-2013-12558), MF is supported by a predoctoral fellowship from the Spanish Ministry of Economy and Competitiveness MINECO (BES-2016-078420), and TS-A is supported by a predoctoral fellowship from the Basque Government.
