**Acknowledgements**

The authors thank the Ministry of Education and Sciences of the Republic of Kazakhstan for funding the project 074/GF "The creation and study of mutant genotypes of wheat for identifying valuable breeding forms and new alleles of genes controlling key adaptive properties" and AP05131881 "Development of integrated approaches for bifortification, high bioavailability of the most important micronutrients of spring wheat and health." The authors wish to acknowledge the technical financial support of the International Atomic Energy Agency (IAEA, Austria) for National TC project KAZ/5003, "Increasing Micronutrient Content and Bioavailability in Wheat Germplasm by Means of an Integrated Approach."
