**Acknowledgements**

This study was supported by the National Key R&D Program of China (2016 YFD0100500, 2016YFD0100300, 2016YFD0100201-21), the National Natural Science Foundation of China (31701449, 31471516, 31401465, 31400074, 31501332), the Natural Science Foundation of Heilongjiang (QC2017013), the Young Innovative Talent training plan of undergraduate colleges and universities in Heilongjiang province (UNPYSCT-2016144), special financial aid to post-doctor research fellow in Heilongjiang (To Qi Zhaoming), the Heilongjiang Funds for Distinguished Young Scientists (JC2016004) and the Outstanding Academic Leaders Projects of Harbin, China (2015RQXXJ018), the China Post Doctoral Project (2015 M581419), the Dongnongxuezhe Project (to Chen Qingshan), the Young Talent Project (to Qi Zhaoming, 518062) of Northeast Agricultural University and SIPT project of Northeast Agriculture University (2018-171, 2018-172).

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