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DOI: 10.1371/journal.pone.0062406

\*Address all correspondence to: h.donoghue@ucl.ac.uk

1 Centre for Clinical Microbiology, University College London, London, UK

2 Department of Microbial Sciences, Faculty of Health and Medical Sciences, University of Surrey, Guildford, UK

3 Mycobacteriology Unit, Department of Biomedical Sciences, Institute of Tropical Medicine, Antwerp, Belgium

4 Institute of Microbiology and Infection, School of Biosciences, University of Birmingham, Birmingham, UK
