**Author details**

Yang Liu1 \*, Jeffrey L. Walck<sup>2</sup> \* and Yousry A. El-Kassaby1 \*

\*Address all correspondence to: yang.liu@ubc.ca, Jeffrey.Walck@mtsu.edu and y.el-kassaby@ubc.ca

1 Department of Forest and Conservation Sciences, University of British Columbia, Vancouver, BC, Canada

2 Department of Biology, Middle Tennessee State University, Murfreesboro, TN, USA
