**Acknowledgements**

The present study was funded by the Research Unit of Bio-ecology and Evolutionary Systematics (UR11ES11), Faculty of Science of Tunis, University of Tunis El Manar. We would like to thank Dr. Dieter Garbe-Schönberg (Institut für Geowissenschaften, ICP-MS Labor, Universität zu Kiel, Kiel, Germany) and Dr. Martin Zimmer (Leibniz Center for Tropical Marine Ecology Fahrenheitstr. Bremen, Bermany) for assistance with soil analysis.
