**Acknowledgements**

We are indebted to Paula Alcazar for her technical assistance. This research was supported by grants CTM/2008-00304/ANT, CTM 2010-12134-E/ANT and CTM2011-16003-E from the Spanish Ministerio de Economía y Competitividad (MINECO). E.G.L. is recipient of a MINECO Fellowship (Grant PTAT2010-03424, Programa Nacional de Contratación e Incorporación de RRHH).
