**Acknowledgements**

This work was supported by the Key Laboratory of Yunnan Province Universities of the Diversity and Ecological Adaptive Evolution for Animals and Plants on Yun-Gui Plateau, the Scientific Research Foundation of Yunnan Provincial Department of Education (2017ZZX186), and the National Natural Science Foundation of China (Nos. NSFC 31760013, NSFC 31260087, NSFC 31460179, and NSFC 31460561). The authors would like to thank the Yunnan Province Universities of the Science and Technology Innovation Team for the exploitation and utilization of endophytes for finance support.
