**Acknowledgements**

This work was supported by Becas Doctorado Nacional CONICYT and funded in part by DAND Codelco-Andina, Fondecyt 1151375 and Fondap-Ideal 15150003 to L.V-C. We are grateful to Loretto Arriagada, Alfonso Jara, Gustavo Diaz, Jorge González, Cindy Cerna, and Aliro Manosalva for technical assistance in the field and laboratory.
