**Author details**

Catherine Marie Breton *Present address: CNRS ISE-M UMR 5554, Montpellier, France Address: INRA , TGU AGAP, Equipe DAVEM, Montpellier, France* 

Peter Warnock *Missouri Valley College, USA* 

André Jean Bervillé *INRA, UMR DIAPC, Montpellier, France* 
