**Author details**

Hideya Takahashi\*

*Ushimado Marine Institute, Faculty of Science, Okayama University, Ushimado, Setouchi, Japan Graduate School of Natural Science and Technology, Okayama University, Kitaku Tsushimanaka, Okayama, Japan* 

*Department of Environmental Science, Faculty of Science, Niigata University, Ikarashi, Niigata, Japan* 

Hiroki Kudose, Chiyo Takagi and Tatsuya Sakamoto

*Ushimado Marine Institute, Faculty of Science, Okayama University, Ushimado, Setouchi, Japan Graduate School of Natural Science and Technology, Okayama University, Kitaku Tsushimanaka, Okayama , Japan* 

Shunsuke Moriyama *School of Marine Biosciences, Kitasato University, Ofunato, Iwate, Japan* 
