**Molecular Biomarkers for Ancient Tuberculosis**

David E. Minnikin1, Oona Y-C. Lee1, Houdini H.T. Wu1,

Gurdyal S. Besra1 and Helen D. Donoghue2  *1School of Biosciences, University of Birmingham, Edgbaston, Birmingham, 2Research Department of Infection and The Centre for the History of Medicine, University College London, UK* 
