**6. Acknowledgment**

We thank Dr. S. Sato and Dr. S. Tabata at the Kazusa DNA Research Institute, for nucleotide sequencing of BAC and TAC clones, Dr. H. Kanamori at the Institute of the Society for Techno-innovation of Agriculture and Dr. Y. Katayose at the National Institute of Agrobiological Sciences for nucleotide sequencing of the *E3* genes, and Dr. R. L. Nelson at the U. S. Department of Agriculture, University of Illinois, for supplying seeds of the Harosoy, Harosoy-*E1,* -*E2* and -*e3*. This study was partly supported by the Programs for the Promotion of Basic Research Activities for Innovative Biosciences, Research Fellowships from the Japan Society for the Promotion of Science for Young Scientists (17-2554), a grant from Ministry of Agriculture, Forestry and Fisheries of Japan (Genomics for Agricultural Innovation, DD-2040), from Grant-in Aid for Scientific Research A (18208001).
