**9. Acknowledgements**

We would like to thank Rich Landa and Rachel Steinhardt for their excellent assistance. Many thanks to Chuanmin Hu, and Steve Lohrenz. We thank our reviewers for their excellent and generous comments on our earlier work. Our work was partially funded by the National Atmospheric and Space Administration's Biodiversity Program 05-TEB/05- 0016, the National Oceanic and Atmospheric Administration (NOAA) and the National Environmental Research Council (NERC) UK.
