**Acknowledgements**

This research was funded by the Research Institute for Humanity and Nature for the project, "Biodiversity-driven Nutrient Cycling and Human Well-being in Social-Ecological Systems (D06-14200119) (Leader: Noboru Okuda)," JSPS KAKENHI Grant-in-Aid for Scientific Research (B) 20H03090 and (C) 21 K12359, and Lake Governance Research Center at the University of Shiga Prefecture.
