**Acknowledgements**

This research was supported by National Natural Science Foundation of China (NSFC) projects (Nos 32171320 and 81371667), Xi'an Association for Science and Technology young Talent Promotion Plan (Nos. 095920211324 and 095920221341), Qinchuangyuan cited the high-level innovation and entrepreneurship talent program (QCYRCXM-2022-27), State Key Laboratory of Veterinary Biotechnology Foundation (SKLVBF202207), the Fundamental Research Funds for the Central Universities (No. QTZX22067), the Opening Project of the State Key Laboratory of Microbial Resources (SKLMR-20220703).
