**Acknowledgements**

This work is supported by the Higher Institution Centre of Excellence (HICOE) Research Grant (Innovative Vaccines and Therapeutics against Fish Diseases) (Project No. 6369100), and SATREPS (JICA-JST): COSMOS-MOHE G4-B Research Grant (Microalgae for Sustainable Aquaculture Health: Microalgae Vaccine Delivery System) (Project No. 6300866).
