**Acknowledgements**

This work was funded by the projects RTA2017-00093-00-00 from the National Institute for Agricultural and Food Research and Technology (INIA) and LMP164\_18 from the Government of Aragón; and by the Operational Programme FEDER Aragón 2014-2020 and the European Social Fund from the European Union [Grupo Consolidado A12-17R: "Grupo de investigación en fruticultura: caracterización, adaptación y mejora genética"]. We gratefully acknowledge the Vegetable Germplasm Bank of Zaragoza (BGHZ-CITA, Spain) for supplying the seeds used for this work. I. M.-L. was granted with a predoctoral contract for training doctors from the Spanish Ministry of Science, Innovation and Universities (MCIU) and the Spanish State Research Agency (AEI).
