**2.2 Transcription factors regulating fatty acid biosynthesis**

*WRINKLED1* (WRI1, AT3G54320) directly controls the transcriptional activation of the fatty acid biosynthetic pathway in *Arabidopsis* and belongs to the APETALA2-ethylene-responsive element-binding protein (AP2-EREBP) family [17]. The ortholog of *AtWRI1* in oil palm was validated as a key transcription factor associated with lipid synthesis [3]. In coconut, three AT3G54320 orthologs were found— CCG005292.1, CCG012597.1, and CCG019741.1. CCG005292.1 and CCG012597.1 were expressed in the endosperm but had low expression in leaf and endosperm, while CCG019741.1 has no expression in leaf, embryo, or endosperm [5]. The *CnWRI1* gene copy (CCG012597.1) validated its interaction with the promoter sequence of acetyl-CoA carboxylase by yeast one-hybrid system [15]. Overexpression of *CnWRI1* (CCG012597.1) specifically in *Arabidopsis* seed showed an increase of palmitic acid (C16:0) and linolenic acid (C18:3) but a decrease in oleic acid content [15].
